BNF = biological nitrogen fixation

N = soil nitrogen availability

C:N = carbon-nitrogen ratio of leaves

G = grazing intensity

B = browsing intensity

Rain = available soil water on a seasonal-annual basis

NPP = herb layer net primary production

L = standing fuel load

F = frequency and intensity of fires.

Savanna complexity affecting savanna structure at landscape-to-patch scales. Box and arrow sizes indicate relative importance of pool or process. Solid arrows indicate primary direction of control and dotted arrows indicate positive feed-back processes that maintain the "fertile" and "infertile" stable states.